Essay On Wild Cats

Not to be confused with Feral cat.

This article is about the Old World wildcat. For other uses, see Wildcat (disambiguation).

The wildcat is a small cat species complex comprising Felis silvestris and the Felis lybica. The former is native to Europe and the Caucasus; the latter to much of Africa, Southwest and Central Asia into India, and western China.[3]

Because of the species' wide range the wildcat is listed as Least Concern on the IUCN Red List since 2002. However, crossbreeding of wildcats and domestic cat (Felis catus) occurs in particular in Europe and is considered a potential threat for the preservation of the wild species.[1]

The wildcat shows a high degree of geographic variation. Whereas the Asiatic wildcat is spotted, the African wildcat is faintly striped, has short sandy-gray fur, banded legs, red-backed ears and a tapering tail. The European wildcat is striped, has long fur and a bushy tail with a rounded tip, and is larger than a domestic cat.[4]

The African wildcat is the ancestor of the domestic cat. Genetic, morphological and archaeological evidence suggests that domestication of Old-World wildcats began approximately 7500 years BCE in the Fertile Crescent region of the Near East. The association of wildcats with humans appears to have developed along with the growth of agricultural villages during the Neolithic Revolution, with wildcats preying on rodents that infested the grain stores of early farmers. Results of a phylogeographic analysis suggest to include the Chinese mountain cat as a Felis silvestris subspecies.[2]

In 2005, 22 wildcat subspecies were recognised as valid taxa.[5]

Taxonomy[edit]

In 1778, Johann von Schreber described the European wildcat using the scientific nameFelis (catus) silvestris.[6] In 1780, Georg Forster described a cat from the Barbary Coast using the name Felis lybica.[7] In subsequent decades, several naturalists and explorers described 40 wildcats from European, African and Asian countries. The taxonomist Reginald Innes Pocock reviewed the collection of wildcat skins in the British Museum, and in 1951 designated seven Felis silvestrissubspecies from Europe to Asia Minor, and 25 Felis lybica subspecies from Africa, and West to Central Asia.[8]

As of 2005[update], 22 subspecies were recognised by Mammal Species of the World.[5] They were divided into three groups:[9][10]

  • Forest wildcats (silvestris group).
  • Steppe wildcats (ornata-caudata group): distinguished from the forest wildcats by their smaller size, longer, more sharply pointed tails, and comparatively lighter fur colour;[11] includes the subspecies ornata, nesterovi and iraki.[12]
  • Bay or bush wildcats (ornata-lybica group): distinguished from the steppe wildcats by their generally paler colouration, well-developed spot patterns and bands; includes the subspecies chutuchta, lybica, ocreata, rubida, cafra, griselda, and mellandi.[12] The domestic cat is thought to have derived from this group.[13][14]

The following tables are based on the classification of the species provided in Mammal Species of the World.[5] They also reflect the classification used in the revision of the Cat Classification Task Force:[3]

Forest wildcat subspeciesCharacteristicsDistribution
F. silvestris silvestrisSchreber, 1777, syn.F. s. ferus Erxleben, 1777; obscura Desmarest, 1820; hybrida J. B. Fischer, 1829; ferox Martorbelli, 1896; morea Trouessart, 1904; grampia Miller, 1907; tartessia Miller, 1907; molisana Altobello, 1921; reyi Lavauden, 1929; jordansi Schwarz, 1930; euxina Pocock, 1943; cretensis Haltenorth, 1953A large subspecies, measuring 40–91 cm in body length, 28–35 cm in tail length, and weighing 3.75–11.5 kg. Its fur is dark, with a gray tone. The pattern on the head, the dorsal band and the transverse stripes and spots on the trunk are distinct and usually vivid.[15]
Most of continental Europe, Scotland, Crete, Balearic Islands, Corsica
F. s. caucasicaSatunin, 1905; trapezia Blackler, 1916Smaller than silvestris, measuring 70–75 cm in body length, 26–28 cm in shoulder height, and weighing usually 5.20–6 kg. Its fur is generally lighter than that of silvestris, and is grayer in shade. The patterns on the head and the dorsal band are well developed, though the transverse bands and spots on the trunk are mostly faint or absent. The tail has a black tip, and only three distinct, black transverse rings.[16]
Caucasus and Asia Minor
Steppe wildcat subspeciesCharacteristicsDistribution
F. lybica lybicaForster, 1780; syn. F. l. ocreata Gmelin, 1791; nubiensis Kerr, 1792; guttata Hermann, 1804; maniculata Cretschmar, 1826; mellandi Schwann, 1904; rubida Schwann, 1904; ugandae Schwann, 1904; nandae Heller, 1913; taitae Heller, 1913; nesterovi Birula, 1916; iraki Cheesman, 1921; hausa Thomas and Hinton, 1921; griselda Thomas, 1926; brockmani Pocock, 1944; foxi Pocock, 1944; pyrrhus Pocock, 1944; gordoni Harrison, 1968This subspecies has palish, buffish or light-grayish fur, and a tinge of red on the dorsal band.[17]
Arabian Peninsula, Mespotamia, Israel, Syria and Palestine, Kuwait, Iraq, southwestern Iran, Ethiopia, Guinea, French Sudan and Nigeria in West Africa[18]East Africa, Sudan and Sahel woodlands, northeastern part of the Congo basin, Angola, southern part of the Congo basin and northern Zimbabwe
F. lybica cafraDesmarest, 1822; syn. F. l. xanthella Thomas, 1926; vernayi Roberts, 1932This subspecies has a pale fur with a faint pattern.[19]
Southern Africa
F. lybica ornataGray, 1830; syn. syriaca Tristam, 1867; caudata Gray, 1874; maniculata Yerbury and Thomas, 1895; griseoflava Zukowsky, 1915; kozlovi Satunin, 1905; matschiei Zukowsky, 1914; longipilis Zukowsky, 1915; schnitnikovi Birula, 1915; macrothrix Zukowsky, 1915; murgabensis Zukowsky, 1915; issikulensis Ognev, 1930; tristrami Pocock, 1944This subspecies has light, ochreous-gray coloured fur with dark spots on the back.[20]
Kazakhstan, Transcaucasia, Iran, Afghanistan, and Dzhungaria, western India

Evolution[edit]

Origins[edit]

The wildcat's direct ancestor was Felis lunensis, or Martelli's wildcat, which lived in Europe as early as the late Pliocene. Fossil remains of the wildcat are common in cave deposits dating from the last ice age and the Holocene.[21] At some point during the Late Pleistocene (possibly 50,000 years ago), the wildcat migrated from Europe into the Middle East, giving rise to the steppe wildcat phenotype. Within possibly 10,000 years, the steppe wildcat spread eastwards into Asia and southwards to Africa.[4]

The wildcat's closest living relatives are the sand cat, the Chinese mountain cat (which may be a subspecies of wildcat), the jungle cat and the black-footed cat.[22] As a whole, the wildcat (along with the jungle and leopard cat) represents a much less specialised form than the sand cat and manul. However, wildcat subspecies of the lybica group do exhibit some further specialisation, namely in the structure of the auditory bullae, which bears similarity to those of the sand cat and manul.[23]

Domestication[edit]

The earliest evidence of wildcat taming comes from 9,500-year-old Neolithic graves excavated in Shillourokambos, Cyprus, that contained the skeletons of a human and a cat, buried close to one another. As no records of native cats in Cyprus exist, this discovery indicates that Neolithic farmers brought cats to Cyprus from the Middle East, most likely to control rodents.[24][25][26][27] Wildcats were probably domesticated in the Fertile Crescent around the time of the introduction of agriculture.[2]

Despite thousands of years of domestication, there is very little difference between the housecat and its wild ancestor, as its breeding has been more subject to natural selection imposed by its environment, rather than artificial selection by humans.[14] The wildcat subspecies that gave rise to the housecat is most likely the African wildcat, based on genetics,[2]morphology,[13][14] and behaviour. The African wildcat lacks the sharply defined dorsal stripe present in the European wildcat, a trait which corresponds with the coat patterns found in striped tabbies. Also, like the African wildcat, the housecat's tail is usually thin, rather than thick and bushy like the European wildcat's.[28] In contrast to European wildcats, which are notoriously difficult to tame,[13][29] hand-reared African wildcats behave almost exactly like domestic tabbies, but are more intolerant of other cats, and almost invariably drive away their siblings, mates, and grown kittens.[30] Further evidence of an African origin for the housecat is present in the African wildcat's growth; like housecat kittens, African wildcat kittens undergo rapid physical development during the first two weeks of life. In contrast, European wildcat kittens develop much more slowly.[31] The bacula of European domestic cats bear closer resemblance to those of local, rather than African wildcats, thus indicating that crossbreeding between housecats and wildcats of European origin has been extensive.[30]

Characteristics[edit]

Body size[edit]

Compared to other members of the Felinae, the wildcat is a small species, but is nonetheless larger than the housecat.[32] The wildcat is similar in appearance to a striped tabby cat, but has relatively longer legs, a more robust build, and a greater cranial volume.[33] The tail is long, and usually slightly exceeds one-half of the animal's body length. Its skull is more spherical in shape than that of the jungle and leopard cat. The ears are moderate in length, and broad at the base. The eyes are large, with vertical pupils, and yellowish-green irises.[32] Its dentition is relatively smaller and weaker than the jungle cat's.[34] The species size varies according to Bergmann's rule, with the largest specimens occurring in cool, northern areas of Europe (such as Scotland and Scandinavia) and of Middle Asia (such as Mongolia, Manchuria and Siberia).[35] Males measure 43 to 91 cm (17 to 36 in) in body length, 23 to 40 cm (9.1 to 15.7 in) in tail length, and normally weigh 5 to 8 kg (11 to 18 lb). Females are slightly smaller, measuring 40 to 77 cm (16 to 30 in) in body length and 18 to 35 cm (7.1 to 13.8 in) in tail length, and weighing 3 to 5 kg (6.6 to 11.0 lb).[34][36] Both sexes possess pre-anal glands, which consist of moderately sized sweat and sebaceous glands around the anal opening. Large-sized sebaceous and scent glands extend along the full length of the tail on the dorsal side. Male wildcats have pre-anal pockets located on the tail, which are activated upon reaching sexual maturity. These pockets play a significant role in reproduction and territorial marking. The species has two thoracic and two abdominal teats.[37] The wildcat has good night vision, having 20 to 100% higher retinal ganglion cell densities[vague] than the housecat. It may[vague] have colour vision as the densities of its cone receptors are more than 100% higher than in the housecat. Its sense of smell is acute, and it can detect meat at up to 200 metres.[22] The wildcat's whiskers are white; they can reach 5 to 8 cm in length on the lips, and number 7 to 16 on each side. The eyelashes range from 5 to 6 cm in length, and can number 6 to 8 per side. Whiskers are also present on the inner surface of the wrist,[dubious– discuss] and can measure 3 to 4 cm.[38]

Fur[edit]

Forest wildcat[edit]

The forest wildcat's fur is fairly uniform in length throughout the body. The hair on the tail is very long and dense, thus making it look furry and thick. In winter, the guard hairs measure 7 cm, the tip hairs 5.5–6 cm, and the underfur 4.5–5.5 cm. Corresponding measurements in the summer are 5–6.7 cm, 4.5–6 cm, and 5.3 cm. In winter, the forest wildcat's main coat colour is fairly light gray, becoming richer along the back, and fading onto the flanks. A slight ochreous shade is visible on the undersides of the flanks. A black and narrow dorsal band starts on the shoulders, and runs along the back, usually terminating at the base of the tail. Indistinct black smudges are present around the dorsal band, which may form a transverse striping pattern on rare occasions. The undersurface of the body is very light gray, with a light ochreous tinge. One or more white spots may occur on rare occasions on the throat, between the forelegs, or in the inguinal region. The tail is the same colour as the back, with the addition of a pure black tip. 2–3 black, transverse rings occur above the tail tip. The dorsal surface of the neck and head are the same colour as that of the trunk, but is lighter gray around the eyes, lips, cheeks, and chin. The top of the head and the forehead bear four well-developed dark bands. These bands sometimes split into small spots which extend to the neck. Two short and narrow stripes are usually present in the shoulder region, in front of the dorsal band. A dark and narrow stripe is present on the outer corner of the eye, under the ear. This stripe may extend into the neck. Another such stripe occurs under the eye, which also extends into the neck. The wildcat's summer coat has a fairly light, pure background colour, with an admixture of ochre or brown. In some animals, the summer coat is ashen coloured. The patterns on the head and neck are as well-developed as those on the tail, though the patterns on the flanks are almost imperceptible.[38]

Steppe wildcat[edit]

The steppe wildcat's coat is lighter than the forest wildcat's, and never attains the level of density, length, or luxuriance as that of the forest wildcat, even in winter. The tail appears much thinner than that of the forest wildcat, as the hairs there are much shorter, and more close-fitting. The colours and patterns of the steppe wildcat vary greatly, though the general background colour of the skin on the body's upper surface is very lightly coloured. The hairs along the spine are usually darker, forming a dark gray, brownish, or ochreous band. Small and rounded spots cover the entirety of the species' upper body. These spots are solid and sharply defined, and do not occur in clusters or appear in rosette patterns. They usually do not form transverse rows or transverse stripes on the trunk, as is the case in the forest wildcat. Only on the thighs are distinct striping patterns visible. The underside is mainly white, with a light gray, creamy or pale yellow tinge. The spots on the chest and abdomen are much larger and more blurred than on the back. The lower neck, throat, neck, and the region between the forelegs are devoid of spots, or have bear them only distinctly. The tail is mostly the same colour as the back, with the addition of a dark and narrow stripe along the upper two-thirds of the tail. The tip of the tail is black, with 2–5 black transverse rings above it. The upper lips and eyelids are light, pale yellow-white. The facial region is of an intense gray colour, while the top of the head is covered with a dark gray coat. In some specimens, the forehead is covered in dense clusters of brown spots. A narrow, dark brown stripe extends from the corner of the eye to the base of the ear.[39]

Behaviour[edit]

Social and territorial behaviours[edit]

The wildcat is a largely solitary animal, except during the breeding period. The size of its home range varies according to terrain, the availability of food, habitat quality, and the age structure of the population. Male and female ranges overlap, though core areas within territories are avoided by other cats. Females tend to be more sedentary than males, as they require an exclusive hunting area when raising kittens.[40] Within its territory, the wildcat leaves scent marks in different sites, the quantity of which increases during estrus, when the cat's preanal glands enlarge and secrete strong smelling substances, including trimethylamine.[41] Territorial marking consists of urinating on trees, vegetation and rocks, and depositing faeces in conspicuous places. The wildcat may also scratch trees, leaving visual markers, and leaving its scent through glands in its paws.[40]

The wildcat does not dig its own burrows, instead sheltering in the hollows of old or fallen trees, rock fissures, and the abandoned nests or earths of other animals (heron nests, and abandoned fox or badger earths in Europe,[42] and abandoned fennec dens in Africa[43]). When threatened, a wildcat with a den will retreat into it, rather than climb trees. When taking residence in a tree hollow, the wildcat selects one low to the ground. Dens in rocks or burrows are lined with dry grasses and bird feathers. Dens in tree hollows usually contain enough sawdust to make lining unnecessary. During flea infestations, the wildcat leaves its den in favour of another. During winter, when snowfall prevents the wildcat from travelling long distances, it remains within its den more than usual.[42]

Hunting behaviour[edit]

When hunting, the wildcat patrols forests and along forest boundaries and glades. In favourable conditions, it will readily feed in fields. The wildcat will pursue prey atop trees, even jumping from one branch to another. On the ground, it lies in wait for prey, then catches it by executing a few leaps, which can span three metres. Sight and hearing are the wildcat's primary senses when hunting, its sense of smell being comparatively weak. When hunting aquatic prey, such as ducks or nutrias, the wildcat waits on trees overhanging the water. It kills small prey by grabbing it in its claws, and piercing the neck or occiput with its fangs. When attacking large prey, the wildcat leaps upon the animal's back, and attempts to bite the neck or carotid. It does not persist in attacking if prey manages to escape it.[44] Wildcats hunting rabbits have been observed to wait above rabbit warrens for their prey to emerge.[45] Although primarily a solitary predator, the wildcat has been known to hunt in pairs or in family groups, with each cat devoted entirely to listening, stalking, or pouncing. While wildcats in Europe will cache their food, such a behaviour has not been observed in their African counterparts.[46]

Reproduction and development[edit]

The wildcat has two estrus periods, one in December–February and another in May–July.[47] Estrus lasts 5–9 days, with a gestation period lasting 60–68 days.[45]Ovulation is induced through copulation. Spermatogenesis occurs throughout the year. During the mating season, males fight viciously,[47] and may congregate around a single female. There are records of male and female wildcats becoming temporarily monogamous. Kittens usually appear in April–May, though some may be born from March–August. Litter size ranges from 1–7 kittens.[45]

Kittens are born blind and helpless, and are covered in a fuzzy coat.[47] At birth, the kittens weigh 65-163 grams, though kittens under 90 grams usually do not survive. They are born with pink paw pads, which blacken at the age of three months, and blue eyes, which turn amber after five months.[45] Their eyes open after 9–12 days, and their incisors erupt after 14–30 days. The kittens' milk teeth are replaced by their permanent dentition at the age of 160–240 days. The kittens start hunting with their mother at the age of 60 days, and will start moving independently after 140–150 days. Lactation lasts 3–4 months, though the kittens will eat meat as early as 1.5 months of age. Sexual maturity is attained at the age of 300 days.[47] Similarly to the housecat, the physical development of African wildcat kittens over the first two weeks of their lives is much faster than that of European wildcats.[31] The kittens are largely fully grown by 10 months, though skeletal growth continues for over 18–19 months. The family dissolves after roughly five months, and the kittens disperse to establish their own territories.[45] The species' maximum life span is 21 years, though it usually only lives up to 13–14 years.[47]

Ecology[edit]

Diet[edit]

Throughout its range, small rodents (mice, voles, and rats) are the wildcat's primary prey, followed by birds (especially ducks and other waterfowl, galliformes, pigeons and passerines), dormice, hares, nutria, and insectivores.[48] Unlike the housecat, the wildcat can consume large fragments of bone without ill-effect.[49] Although it kills insectivores, such as moles and shrews, it rarely eats them[48] because of the pungent scent glands on their flanks.[50] When living close to human habitations, the wildcat can be a serious poultry predator.[48] In the wild, the wildcat consumes up to 600 grams of food daily.[51]

The diet of wildcats in Great Britain varies geographically; in eastern Scotland, lagomorphs make up 70% of their diet, while in the west, 47% consists of small rodents.[45] In Western Europe, the wildcat feeds on hamsters, brown rats, dormice, water voles, voles, and wood mice. From time to time, small carnivores (martens, polecats, stoats, and weasels) are preyed upon, as well as the fawns of red deer, roe deer, and chamois. In the Carpathians, the wildcat feeds primarily on yellow-necked mice, red-backed voles, and ground voles. European hares are also taken on occasion. In Transcarpathia, the wildcat's diet consists of mouse-like rodents, galliform birds, and squirrels. Wildcats in the Dnestr swamps feed on small voles, water voles, and birds, while those living in the Prut swamps primarily target water voles, brown rats, and muskrats. Birds taken by Prut wildcats include warblers, ferruginous ducks, coots, spotted crakes, and gadwalls. In Moldavia, the wildcat's winter diet consists primarily of rodents, while birds, fish, and crayfish are eaten in summer. Brown rats and water voles, as well as muskrats and waterfowl are the main sources of food for wildcats in the Kuban delta. Wildcats in the northern Caucasus feed on mouse-like rodents and edible dormice, as well as birds on rare occasions. On rare occasions, young chamois and roe deer, are also attacked. Wildcats on the Black Sea coast are thought to feed on small birds, shrews, and hares. On one occasion, the feathers of a white-tailed eagle and the skull of a kid were found at a den site.[48] In Transcaucasia, the wildcat's diet consists of gerbils, voles, birds, and reptiles in the summer, and birds, mouse-like rodents, and hares in winter. Turkmenian wildcats feed on great and red-tailed gerbils, Afghan voles, thin-toed ground squirrels, Tolai hares, small birds (particularly larks), lizards, beetles, and grasshoppers. Near Repetek, the wildcat is responsible for destroying over 50% of nests made by desert finches, streaked scrub warblers, red-tailed warblers, and turtledoves. In the Qarshi steppes of Uzbekistan, the wildcat's prey, in descending order of preference, includes great and red-tailed gerbils, jerboas, other rodents and passerine birds, reptiles, and insects. Wilcats in eastern Kyzyl Kum have similar prey preferences, with the addition of tolai hares, midday gerbils, five-toed jerboas, and steppe agamas. In Kyrgyzstan, the wildcat's primary prey varies from tolai hares near Issyk Kul, pheasants in the Chu and Talas valleys, and mouse-like rodents and gray partridges in the foothills. In Kazakhstan's lower Ili, the wildcat mainly targets rodents, muskrats, and Tamarisk gerbils. Occasionally, remains of young roe deer and wild boar are present in its faeces. After rodents, birds follow in importanance, along with reptiles, fish, insects, eggs, grass stalks and nuts (which probably enter the cat's stomach through pheasant crops).[52] In west Africa, the wildcat feeds on rats, mice, gerbils, hares, small to medium-sized birds (up to francolins), and lizards. In southern Africa, where wildcats attain greater sizes than their western counterparts, antelope fawns and domestic stock, such as lambs and kids are occasionally targeted.[43]

Predators and competitors[edit]

Because of its habit of living in areas with rocks and tall trees for refuge, dense thickets and abandoned burrows, wildcats have few natural predators. In Central Europe, many kittens are killed by pine martens, and there is at least one account of an adult wildcat being killed and eaten.[53] In the steppe regions of Europe and Asia, village dogs constitute serious enemies of wildcats, along with the much larger Eurasian Lynx, one of the rare habitual predators of healthy adults. In Tajikistan, wolves are their most serious enemies, having been observed to destroy cat burrows. Birds of prey, including eagle-owls and saker falcons, have been known to kill wildcat kittens.[54]Seton Gordon recorded an instance where a wildcat fought a golden eagle, resulting in the deaths of both combatants.[55] In Africa, wildcats are occasionally eaten by pythons.[56] Competitors of the wildcat include the jungle cat, golden jackal, red fox, marten, and other predators. Although the wildcat and the jungle cat occupy the same ecological niche, the two rarely encounter one another, on account of different habitat preferences: jungle cats mainly reside in lowland areas, while wildcats prefer higher elevations in beech forests.[53]

Communication[edit]

The wildcat is a mostly silent animal.[40] The voice of steppe wildcats differs little from the housecat's, while that of forest wildcats is similar, but coarser.[57]

Name/TranscriptionSound descriptionPostureContext
BrrroooA rolling turtledove-like call.[58]Emitted as a greeting and as a means of self-identification.[58]
Hiss
MauSimilar to a housecat's miaow, but with the preliminary ee omitted.[59]Emitted by kittens requesting food.[59]
Meeeoo! Meeeoo!A piercing buzzard-like call that can be heard 200 yards away.[60]Distress call emitted by kittens.[60]
Noine, noine, noineEmitted by adults feeding contentedly.[61]
PAAAH!Accompanied by bracing and stamping of forelimbs.[62]Emitted when angered.[62]
RumbleTranscribed as urrr urrr, and described by Mike Tomkies as sounding "like a dynamo throbbing deep in the bowels of the earth".[63]Emitted when approached by humans, but does not attack.[64]
SquawkA loud squawking noise, similar to that of ducks.[65]Emitted by kittens grabbed by the scruff of the neck.[65]
Wheeou wheeouA high pitched whistle, similar to a weak buzzard call. The sound is piercing, but not far-carrying.[29]Made with the mouth barely open.[29]Emitted by kittens summoning their mother.[29]

Diseases and parasites[edit]

The wildcat is highly parasitised by helminths. Some wildcats in Georgia may carry five helminth species: Hydatigera taeniaeformis, Diphyllobothrium mansoni, Toxocara mystax, Capillaria feliscati and Ancylostoma caninum. Wildcats in Azerbaijan carry Hydatigera krepkogorski and T. mystax. In Transcaucasia, the majority of wildcats are infested by the tickIxodes ricinus. In some summers, wildcats are infested with fleas of the Ceratophyllus genus, which they likely contract from brown rats.[53]

Distribution[edit]

The wildcat's distribution is very broad, encompassing most of Africa, Europe, and southwest and central Asia into India, China, and Mongolia. Subspecies are distributed as follows:[1]

  • The African wildcat (F. s. lybica) occurs across northern Africa, around the Arabian Peninsula's periphery to the Caspian Sea, encompassing a wide range of habitats, with the exception of closed tropical forests. It occurs throughout the savannahs of West Africa, from Mauritania on the Atlantic seaboard eastwards to the Horn of Africa (Somalia, Ethiopia, Eritrea, and Djibouti) and Sudan. In north Africa, it occurs discontinuously from Morocco through Algeria, Tunisia, Libya into Egypt. Small numbers occur in true deserts such as the Sahara, particularly in hilly and mountainous areas, such as the Hoggar.
  • The Southern African wildcat (F. s. cafra) is distributed in all east and southern African countries. The border between the two subspecies is estimated to occur in the area of Tanzania and Mozambique.
  • The Asiatic wildcat (F. s. ornata) ranges from the east of the Caspian Sea into western India, north to Kazakhstan and into western China and southern Mongolia.
  • The Chinese mountain cat (F. s. bieti) is indigenous to western China, and is particularly abundant in the Qinghai and possibly Sichuan provinces.
  • The European wildcat (F. s. silvestris) was once very widely distributed in Europe and absent only in Fennoscandia and Estonia. Between the late 1700s and mid-1900s, it was extirpated locally so that its European range became fragmented. In the Pyrenees, it occurs from sea level to 2,250 m (7,380 ft). It is possible that in some areas, including Scotland and Stromberg, Germany, pure wildcats have crossbred extensively with domestic cats. The only islands in the Mediterranean with native populations of wildcats are Sicily, Sardinia, Corsica and possibly Crete, where wildcats likely descended from feral populations introduced in Neolithic times. It is possibly extinct in the Czech Republic, and considered regionally extinct in Austria, though vagrants from Italy are spreading into Austrian territory.

The European wildcat was thought extinct in the Netherlands.[1] In 2006, a wildcat was photographed by a camera trap in the province of Limburg. Since then there were frequent, but unconfirmed sightings in this province until December 2012 when a cat was photographed again. A male wildcat was photographed several times in April 2013 while it was scavenging the carcass of a dead deer, an unusual behavior for a wildcat.[66]

Relationships with humans[edit]

In culture[edit]

In mythology[edit]

In Celtic mythology, the wildcat was associated with rites of divination and Otherworldly encounters. Domestic cats are not prominent in Insular Celtic tradition (as housecats were not introduced to the British Isles until the Mediaeval period).[citation needed] Fables of the Cat Sìth, a fairy creature described as resembling a large white-chested black cat, are thought to have been inspired by the Kellas cat, itself thought to be a free ranging wildcat-houscat crossbreed.[67] Doctor William Salmon, writing in 1693, mentioned how portions of the wildcat were used for medicinal purposes; its flesh was used to treat gout, its fat used for dissolving tumours and easing pain, its blood used for curing "falling sickness", and its excrement used for treating baldness.[68]

In heraldry[edit]

The wildcat is considered an icon of the Scottish wilderness, and has been used in clan heraldry since the 13th century.[67] The Picts venerated wildcats, having probably named Caithness (Land of the Cats) after them. According to the foundation myth of the Catti tribe, their ancestors were attacked by wildcats upon landing in Scotland. Their ferocity impressed the Catti so much, that the wildcat became their symbol.[69] A thousand years later, the progenitors of Clan Sutherland, equally impressed, adopted the wildcat on their family crest.[69][70] The Chief of Clan Sutherland bears the title Morair Chat (Great Man of the Cats). The Clan Chattan Association (also known as the Clan of Cats) is made up of 12 different clans, the majority of which display the wildcat on their badges.[67][70]

In literature[edit]

Shakespeare referenced the wildcat three times:[68]

The patch is kind enough ; but a huge feeder
Snail-slow in profit, and he sleeps by day
More than the wild cat.

— The Merchant of Venice Act 2 Scene 5 lines 47–49

Thou must be married to no man but me ;
For I am he, am born to tame you, Kate ;
And bring you from a wild cat to a Kate
Comfortable, as other household Kates.

— The Taming of the Shrew Act 2 Scene 1 lines 265–268

Thrice the brinded cat hath mew'd.

— Macbeth Act 4 Scene 1 line 1

Hunting[edit]

Although a furbearer, the wildcat's skin is of little commercial value,[57] due to the unattractive colour of its natural state, and the difficulties present in dyeing it.[71] In the former Soviet Union, the fur of a forest wildcat usually fetched 50 kopecks, while that of a steppe wildcat fetched 60 kopecks.[57] Wildcat skin is almost solely used for making cheap scarfs, muffs,[71] and women's coats. It is sometimes converted into imitation sealskin.[57] As a rule, wildcat fur is difficult to dye in dark brown or black, and has a tendency to turn green when the dye is not well settled into the hair. When dye is overly applied, wildcat fur is highly susceptible to singeing.[71]

In the former Soviet Union, wildcats were usually caught accidentally in traps set for martens. In modern times, they are caught in unbaited traps on pathways or at abandoned fox, badger, hare or pheasant trails. One method of catching wildcats consists of using a modified muskrat trap with a spring placed in a concealed pit. A scent trail of pheasant viscera leads the cat to the pit.[57] A wildcat caught in a trap growls and snorts.[72]

References[edit]

Skulls of a wildcat (top left), a housecat (top right), and a hybrid between the two (bottom centre)
Skin of a European forest wildcat
Skin of an Indian steppe wildcat
European wildcat killing a deer fawn, as illustrated in Lydekker's Wild Life of the World (1916)

“A Safe Cat is a Happy Cat”

They are playful and loving, aloof and mysterious, and frisky and mischievous. They are also becoming the most frequent occupants of America’s animal shelters, where millions of them are cared for each year.

They are cats – America’s most popular pets but also the pets most likely to die prematurely from disease, poisons, attacks by other animals, abuse by humans, or speeding vehicles.

Cats are deserving of our protection as dogs. But millions of cats suffer and die needlessly because they are allowed to roam. The vast majority of these cats are not the victims of cruel or thoughtless owners; in fact, their caregivers often love them like children. Instead cats are the victims of outmoded perceptions that cast them as independent, natural explorers who prefer to be left to their own devices.

The Myth of The Outdoor Cat

The good news is that cats do not need to wander to lead fulfilling lives. The bad news is that many caregivers believe the opposite. Free-roaming cats get a dangerous tradeoff: freedom to roam in exchange for the vastly increased likelihood of a premature, painful death.

The Humane Society of the United States estimates the average lifespan of a free-roaming cat is less than three years, compared to 12-15 years for the average indoor-only cat. Even cats in suburban neighborhoods can meet untimely fates and never return home.

Safely confined cats avoid these hazards:

 
  • Traffic 

    Collisions with cars and other vehicles are common killers. It is a myth that cats are “streetwise” about cars. Cats are intelligent and alert but, like most other animals, stand little chance against fast-moving vehicles.

 

  • Diseases

    Rabies and other diseases that can be transmitted to humans are a serious public health concern. According to the American Association of Feline Practitioners, as many as 15 percent of sick cats are infected with feline leukemia. This virus is fatal and is transmitted through contact with other cats.

 

  • Poisons

    Poisons exist on chemically treated lawns, in bait left out to kill rats or mice, and in auto antifreeze.

 

  • Other Animals

    Other cats, dogs, and wild predators such as coyotes, raccoons, and foxes are potential enemies of cats and often engage in fights that leave cats injured or dead. Outdoor cats can suffer torn ears, cut eyes, abscesses, and other injuries requiring expensive veterinary treatment.

 

  • Cruel People

    Many shelter workers see cats who have been burned, poisoned, or otherwise tortured by disturbed children and adults.

Other Dangers Lurking Outdoors

Free-Roaming cats inevitably pick up fleas and ticks and then bring these pests into the home. Fleas can cause anemia, skin irritations, and allergies in cats and transmit diseases to humans through their bites. Unsterilized cats allowed to roam contribute to the high number of cats who end up in our nation’s animal shelters every day.

Most veterinarians treat the injuries and diseases resulting from allowing cats outdoors unsupervised. In fact, two out of three veterinarians recommend keeping cats indoors, most often citing dangers from vehicles and disease.

The Myth of the Indoor-Only Cat

Keeping cats safely confined is not new to many long-term cat lovers. But it is news to many people who grew up with indoor-outdoor or outdoor-only felines.

Some cat owners believe that it is unnatural, or even cruel, to keep cats cooped up inside all the time. Unfortunately, this belief is self-perpetuating, especially if the pet caregiver makes no effort to provide the cat with a stimulating indoor environment.

While most cats enjoy being outside where they can hunt prey and explore their surroundings, it’s a myth that going outside is a prerequisite for feline happiness. Playing with an indoor cat easily satisfies the animal’s stalking instinct and keeps the cat stimulated and healthy through exercise. In fact, the indoor cat who gets lots of attention and playtime is happier than the outdoor-indoor cat who is generally ignored by human companions.

Cat owners can easily create feline-friendly homes that meet all their cats’ needs. Many innovative and fun toys can help make the indoor cat life a great but safe adventure.

Cats don’t have to be deprived of the great outdoors to stay safe. Cats can be trained to accept a harness and leash, and cat enclosures can allow them to experience all the pleasures of the great outdoors without all of the risks.

Keeping Communities Safe

Cats allowed to roam freely outside not only face potential harm but also have an unintended impact on our communities.

Local governments spend hundreds of millions of dollars every year controlling stray animals, from neighborhood cats defecating in sandboxes to feral (wild) cats. Cats are now a major focus of local animal care and control agencies, which in the past concentrated most of their efforts on dog control and rescue.

In fact, animal control agencies were established long ago primarily to control the spread of rabies among dogs in the street. Today cats who roam, particularly after dark, are likely to come into contact with nocturnal creatures, including raccoons and skunks, the primary vector species of rabies in the wild. As a result, cats are now the most common domestic vector of rabies, with 278 cases reported in 1999 in the United States.

In addition, free-roaming cats kill millions of wild animals each year. Studies have shown that most of the animals killed are small mammals; approximately 25 percent are birds. Well-fed housecats kill wildlife because of their instinct to hunt prey, not because they need the food. Cats are not a part of natural ecosystems, and their predation causes unnecessary suffering and death to wild animals. In addition they also cause conflict amongst neighbors.

This information is provided courtesy of The Humane Society of the United States and the Safe Cats Campaign.Visit the Humane Society Online.

 

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